Some rituals, like the sacred hunt performed by Shadow Bones, allow the werewolfs to attack a spirit in twilight as if it is materialized.
|World||State of spirit||Damage from bane||Physical damage|
I think in some places it should say materialized instead of manifested. When they are in twilight they are invisible and untouchable by anything that’s not a spirit, but the spirit senses of the werewolf allow them to sense it. I understand it as the werewolf being able to see and hear it but not touch it.